Hakea stenophylla subsp. notialis R.M.Barker, Nuytsia 12: 7 (1998)
T: Murchison sand plain, 320 mile post, North West Coastal Hwy, W.A., 20 July 1972, G.Phillips 27; holo: PERTH; iso: CANB, DNA, K, MEL, NSW, PERTH.
Erect shrub or small tree, 2–4 m high, probably without lignotuber; bark dark, ±smooth. Branchlets densely appressed-pubescent; hairs predominantly white, some ferruginous, quickly glabrescent. Leaves flat, linear to narrowly obovate, 6–12 cm long, 1–4 mm wide, initially densely appressed ferruginous-pubescent, quickly glabrescent; apex abruptly short-acuminate and uncinate.
Peduncle always 2- or 3-branched, 2–9 mm long, densely white-pubescent; rachis 1–1.5 mm long, white-pubescent. Flowers 10–16, white; pedicels 4–6 mm long, densely white appressed-pubescent. Perianth 1.6–2.5 (?–4) mm long, white-pubescent, recurved to claw. Pistil 7–13 mm long; style recurved, rarely looped; pollen presenter oblique.
Fruit obliquely obovate or elliptic, 3–4.5 cm long, 2–2.6 cm wide, abruptly attenuate into an erect apiculum or an apiculum recurved toward the red-brown wood zone (except Smith 1682); valve with red-brown wood zone 2.5–4 mm wide and pale wood zone 6–10 mm wide; beak black. Seed 17–29 mm long, 9.5–16 mm wide, covering c. half valve surface; wing entirely down one side of seed body, partly or fully down other.
Distribution and ecology
Occurs from Carnarvon to Murchison R., W.A., in sand plain or heath.
To plot an up to date distribution map based on herbarium collections for this species see Australia's Virtual Herbarium. Localities outside the native range may represent cultivated or naturalised records.
Derivation of name
From notialis, Latin for southern, a reference to distribution of this species to the south of ssp. stenophylla.
How the infraspecific taxa differ
At present ssp. stenophylla and ssp. notialis can only be definitely distinguished with fruiting material, but further collections and field observations should confirm both habit and habitat differences, with subsp. stenophylla likely to be lignotuberous.
subsp. notialis ssp. stenophylla Fruiting valve: red-brown wood layer width 2.5-4 mm 1.5-2.5 mm Fruiting valve: pale wood width 8-10 mm 3-6 mm Fruiting valve: seed coverage of surface c. 1/2 c. 3/4 Habit Spreading shrub, up to 2 m high Erect shrub or small tree, to 4 m high Lignotuber Present? Absent?
Fruiting valve: red-brown wood layer width
Fruiting valve: pale wood width
Fruiting valve: seed coverage of surface
Spreading shrub, up to 2 m high
Erect shrub or small tree, to 4 m high
Part of the Pedunculata Group of Barker et al. (1999), this group of 4 species (H. arborescens, H. pedunculata, H. persiehana and H. stenophylla) is held together morphologically by the possession of a bare peduncle below the rachis. They also have obscurely-veined leaves, oblique or conical pollen presenters, hairy perianths and woody fruits without horns.
While Bentham referred them to the Grevilleoides, presumably because they may sometimes have a corky bark, it seems likely that these will eventually prove to be an artificial group, with H. arborescens and H. persiehana being closely related but probably not close to H. pedunculata and H. stenophylla. Molecular studies by Mast (unpubl.) have already shown the former but the latter two species were not included in the study.
W.A.: 35 km NE of Kalbarri, 19 May 1968, J.Bannister s.n. (PERTH); 20 km W of Hamelin HS, T.J.Hawkeswood 59 (PERTH); Tamala Stn, May 1960, R.W.Vollprecht s.n. (PERTH).
Link to FloraBase treatment of this species for WA.
I. Holliday, Hakeas. A Field and Garden Guide 194-5 (2005)
J.A..Young, Hakeas of Western Australia. A Field and Identification Guide 108 (2006).