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Electronic Flora of South Australia
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Electronic Flora of South Australia Family Fact Sheet


Phylum Chlorophyta – Order Cladophorales

Thallus reticulate, essentially monostromatic, stipitate or not, attached by rhizoids from basal segments or the stipe, flabellate or ovate or becoming irregular with an obscure base. Blade formed by anastomoses of un-modified or modified segments (tenacula), with the network remaining open or becoming largely filled in by development of small lateral cells. Cells multinucleate, dividing by ingrowth from the lateral walls. Homoplastic, chloroplast reticulate with numerous pyrenoids.

Life history of isomorphic generations, with biflagellate anisogametes and quadriflagellate zoospores.

Taxonomic notes: The three genera described below were earlier placed in the Siphonocladales, but several species have been shown to have centripetal cross wall formation and not segregative division. Some authors (e.g. Egerod 1952, p. 332) maintain two families on the basis of the presence or not of tenacula connecting lateral branches to other branches and the development of juvenile stages. On this basis, Microdictyon and Anadyomene belong to the Anadyomenaceae and Struvea to the Boodleaceae (with tenacula). Such differences are of generic significance but doubtfully of family significance.


EGEROD, L.E. (1952). An analysis of the siphonous Chlorophycophyta. Univ. Calif. Publ. Bot. 25, 325–454.

The Marine Benthic Flora of Southern Australia Part I complete list of references.

Author: H.B.S. Womersley

Publication: Womersley, H.B.S. (31 May, 1984)
The Marine Benthic Flora of Southern Australia
Part I
©Board of the Botanic Gardens and State Herbarium, Government of South Australia


1. Thallus with obscure base, becoming irregular; blade network open, meshes irregular


1. Thallus more or less stipitate, flabellate or ovate, attached by rhizoids from the stipe cell (s); blade network open or largely closed with small cells


2. Thallus flabellate, blade network largely closed with small cells, stipe cells not markedly larger than cells of the veins, tenaculae absent


2. Thallus ovate and stipitate, blade network open, stipe cell much larger than upper cells, 2–8 cm long, tenacula present


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