Family: Amaranthaceae
Halosarcia
Citation:
Paul G. Wilson, Paul G. Wilson, Nuytsia 3:28 (1980).
Derivation: Greek halos, salt; sarx, flesh; referring to the saline habitat and succulent nature of the plants.
Synonymy: Not Applicable Common name: Samphires.
Description:
Glabrous shrubs or herbs; branches when young of globose to cylindrical internodes (segments or articles); segments succulent, the outer portion eventually shrivelling and deciduous, apex shortly cup-shaped or bilobed representing the reduced opposite leaves.
Inflorescence a spike-like thyrse usually terminal to a branchlet, consisting of triads of flowers (cymes) in the axil of each bract (in H. pluriflora up to 7 flowers in a cyme); opposite bracts united or rarely free, succulent; flowers sessile, usually bisexual, in some variants only female; perianth of fused segments, initially membranous or succulent; lobes 3 (2 lateral and a medial abaxial); stamen solitary, abaxial; ovary vertical, membranous or succulent; style slender, 2- or rarely 3-lobed.
Fruiting perianth membranous, succulent, spongy, pithy, crustaceous or horny; seed ovoid to circular; testa membranous to crustaceous; embryo curved; perisperm abundant, lateral.
Distribution:
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Endemic to Australia except for H. indica which also occurs in Malaysia and in countries bordering the Indian Ocean. (P. G. Wilson (1980) Nuytsia 3:28-69.)
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Biology:
No text
Key to Species:
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1. Opposite bracts free from each other |
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2. Seed pale-brown, tuberculate over the embryo |
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H. flabelliformis 2. |
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2. Seed white with prominent uneven scale-like ribs |
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H. pterygosperma 12. |
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1. Opposite bracts united |
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3. Perianth apex rounded to truncate |
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4. Seed white, with scale-like ribs |
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5. Spikes long and slender, even in outline; ribs smooth, only on the circumference of the seed |
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H. lepidosperma 6. |
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5. Spikes short, undulate in outline; ribs uneven, covering the whole of the seed |
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H. pterygosperma 12. |
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4. Seed brown to black, tuberculate to granular |
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6. Seed a dark reddish-brown to black, covered with concentric ribs |
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H. pergranulata 9. |
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6. Seed brown, variously tuberculate |
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7. Bracts denticulate or ciliolate; cymules 3-7-flowered |
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8. Cymules 3-flowered; seed concentrically tuberculate over the embryo |
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H. halocnemoides 4. |
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8. Cymules 3-7-flowered; seed concentrically ridged over the embryo |
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H. pluriflora 10. |
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7. Bracts entire; cymules 3-flowered |
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9. Branches and spikes very slender, smooth in outline (resembling Casuarinaceae branchlets); fruit apex mammillate |
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H. lylei 7. |
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9. Branches undulate in outline |
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10. Fruitlets falling entire (not tearing at the base to expose the seed); abaxial perianth lobe external . |
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H. fontinalis 3. |
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10. Fruitlets tearing at the base to expose the seed; abaxial perianth lobe internal |
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H. halocnemoides 4. |
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3. Perianth dorsiventrally flattened at the apex |
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11. Margins of articles and of bracts denticulate or ciliate |
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12. Perianth in fruit pithy; pericarp horny all over; fruitlets entire (not torn at the base) |
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H. indica 5. |
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12. Perianth thin in fruit, membranous or cartilaginous; pericarp membranous (at least towards the base); fruitlets tearing at the base to expose the seed |
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H. nitida 8. |
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11. Margins of articles and bracts entire |
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13. Perianth pithy in fruit; pericarp horny (at least towards the apex) |
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H. indica 5. |
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13. Perianth various (but not pithy) in fruit |
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14. Perianth papery or soft; fruitlets free from each other |
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H. cupuliformis 1. |
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14. Perianth crustaceous; fruitlets free or united |
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H. syncarpa 13. |
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16. Bracts truncate or undulate on the margin, cup-shaped at the fruiting stage; fruitlets muricate; spikes terminal, pedunculate |
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H. pruinosa 11. |
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16. Bracts undulate, shrivelled at the fruiting stage; spikes usually sessile |
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H. undulata 14. |
Author:
Not yet available
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