Gramineae
Alternative names: Not Applicable
Description:
Herbs "grasses" with alternate leaves embracing the stem with long sheaths which are slit at maturity on the side opposite to the blade and are usually convolute (i.e. 1 margin rolled over the other); sheath ending at the base of the blade in a tongue-like membrane (ligule), sometimes very short or reduced to hairs; stems usually hollow between the nodes; when the blades are narrow the margins usually involute; most of the stem (or sometimes all of it) usually concealed by the leaf sheaths (when they are termed intravaginal innovations); the nodes are the swollen bases of the sheaths.
Flowers small, bisexual, rarely unisexual, naked in the axil of a concave bract (the lemma), opposite to and enclosed in which is another usually 2-nerved bract (the palea); the palea envelopes the flower, which consists of a 1-celled carpel with I ovule and 2 styles with feathery stigmas and usually 3 stamens with versatile anthers; (the term "flower" is generally understood to include the lemma and palea); at the base of the ovary are 2 (rarely 1, 3 or 0) minute delicate scales (the lodicules) usually regarded as the perianth; below the lemma and palea, and enclosing them more or less, are 2 empty bracts (the glumes); all these together form a 1-flowered spikelet, but there may be several flowers arranged alternately on opposite sides of a common axis (the rhachilla)--so that the upper flowers often appear to be stalked-with the glumes at the base, the whole forming a several-flowered spikelet; spikelets arranged in a spike, raceme or panicle.
The fruit (grain or caryopsis) consists of the single albuminous seed and the thin pericarp, which are usually inseparable; at the base of the fruit and on its anterior or outer side (i.e. that which faces the lemma) is a small protuberance showing the position of the embryo; on the posterior or inner side (that facing the palea) is either a punctiform or linear mark (the hilum), which indicates the spot where the ovule was attached to the wall of the ovary; the hilum often lies in a furrow.
Distribution:
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More than 600 genera and 10,000 species, cosmopolitan, more than 200 genera and 1,000 species in Australia.
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Biology:
No text
Taxonomic notes:
Genera are grouped into tribes to bring related genera together. The allocation of genera to tribes is, however, controversial and many of the placings used here may prove to be unsatisfactory.
Key to Genera:
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6. Glumes 5-7-nerved, longer than the lemmas |
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SCHISMUS 15. |
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6. Glumes 1-3-nerved, shorter than the lemmas |
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7. Lemma obtuse or truncate, minutely denticulate |
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PUCCINELLIA 37. |
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7. Lemma notched, often with a short mucro in the notch |
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MOLINERIELLA 50. |
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8. Spikelets 3-10-flowered |
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9. Lemmas rounded or flat on their back |
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10. Perennials; lemmas mucronate |
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DIPLACHNE 70. |
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10. Annuals; lemmas without a mucro |
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DESMAZERIA 29. |
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9. Lemmas keeled by a prominent nerve on their back |
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11. Lemma thin, hairy towards the base |
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POA 36. |
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11. Lemma cartilaginous, glabrous |
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SCLEROCHLOA 38. |
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ISACHNE 18. |
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POA 36. |
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15. Spikelets drooping and quivering |
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BRIZA 26. |
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ERIACHNE 11. |
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12. Spikelet 3-several-flowered |
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16. Lemmas rounded on the back |
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17. Spikelets drooping and quivering |
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BRIZA 26. |
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FESTUCA 31. |
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20. Panicle 1-sided; annuals |
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DESMAZERIA 29. |
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20. Panicle symmetrical; mostly perennials |
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PUCCINELLIA 37. |
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GLYCERIA 32. |
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21. Spikelets sessile, in scattered spikes |
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LEPTOCHLOA 74. |
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21. Spikelets in a loose or contracted panicle |
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22. Spikelets in compact clusters |
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DACTYLIS 28. |
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22. Spikelets not clustered |
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24. Lemma hard, glabrous, 5-nerved; panicle spike- like |
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SCLEROCHLOA 38. |
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24. Lemma thin; panicle usually loose |
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25. Lemma rounded on the back below |
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FESTUCA 31. |
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25. Lemma keeled on the back to the base |
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26. Lemma woolly near the base, 5-9-nerved |
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POA 36. |
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26. Lemma glabrous, 3-nerved |
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SPHENOPUS 39. |
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23. Ligule of hairs or a hair-fringed membrane |
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27. Glumes 1-nerved; lemmas 3-nerved |
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ERAGROSTIS 73. |
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27. Glumes 4-7-nerved; lemmas 7-9-nerved |
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PLAGIOCHLOA 35. |
Author:
Prepared by J.P. Jessop except where indicated
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